Käferinventar von Vorarlberg und Liechtenstein

Transkript

1 Als ehem. Print ISBN Druck & Bindung: Vigl, A-6850 Dornbirn. Eigentümer, Herausgeber und Verleger: Eigenverlag des ERSTEN VORARLBERGER COLEOPTEROLOGISCHEN VEREINS c/o Clemens M. Brandstetter Schesastrasse 1 A-6706 Bürs. Vertrieb in Deutschland: Antiquariat Goecke & Evers, Inh. Erich Bauer Sportplatzweg 5, D Keltern - Weiler. Käferinventar von Vorarlberg und Liechtenstein Insecta: Coleoptera CLEMENS M. BRANDSTETTER & ANDREAS KAPP 1998

2 Voltaire meinte vor ein paar Jahrhunderten, dass die nützlichsten Bücher diejenigen seien, welche den Leser zu ihrer Ergänzung auffordern. Wir entschuldigen uns an dieser Stelle bei jenen Autoren, die wir - ohne irgendwelche Absicht - übersehen haben und bitten diese, uns Ihre Publikationen über Vorarlberg und das Fürstentum Liechtenstein zu melden. Es tut uns leid, hier nicht alle in Vorarlberg und Liechtenstein tatsächlich vorkommenden Käfer präsentieren zu können, denn sicherlich haben es einige bisher geschickt verstanden, sich vor uns zu verbergen. Uns ist aber kein Tal zu weit und kein Berg zu hoch; irgendwann finden wir auch diese Arten, wenn nicht, verweisen wir unsere Nachfolger auf Voltaire. Nicht nach Fundmeldungen durchgeackert wurden: CFA, FHL (14 Bände), HORION (Die Käfer Mitteleuropas). Die von der 'Vorarlberger Naturschau' (KLESS 1974) in Auftrag gegebene Publikation musste grösstenteils ignoriert werden, da sie nicht durch eine körperliche Inventur, sondern anhand von im Museum erstellten Listen erfolgte; jahrelang entstand dadurch der Eindruck, die Müller sche Sammlung samt der Holotype des Lathrobium mülleri befände sich in Dornbirn. Wir wissen einerseits, dass wir mit dieser Publikation einen Teil der Entdeckungsund Forschungsspannung vorwegnehmen, andererseits sind über 60 Publikationen, die Käfer aus Vorarlberg und Liechtenstein betreffen, ein Grund, Inventur zu machen und für die beiden Untersuchungsgebiete auch bisher unveröffentlichte Arten, die allerdings in unserer Umwelt-Datenbank COLDAT gespeichert sind, zu melden. Es sind dies für Vorarlberg 539 und Liechtenstein 456 Arten und wir meinen, dass dies nach so wenigen Jahren der Forschung ein bemerkenswerter Erfolg ist, zumal uns MÜLLER (1912, 1926) mit 3077 Arten für beide Untersuchungsgebiete eine recht stolze Vorgabe hingelegt hat. Wir können anhand der Müller schen Publikationen bestätigen, dass Voltaire wirklich recht hat. Am Anfang und Ende dieses Heftes finden Sie ausklappbare Seiten, sodass Sie diesen die Numerierung der Publikationen und die verwendeten Symbole entnehmen können. Der von manchen Autoren verwendete LUCHT-Code erwies sich bei dieser Inventur als äusserst hilfreich und er sollte zwingender Bestandteil aller faunistischen Arbeiten werden. Vielleicht ist der Tag nicht fern, und die Arten des Planeten Erde erhalten einen international geltenden Strich-Code. Die Nomenklatur sollte auf dem neuesten Stand sein (sensu FHL 1-14). Für Fehlermeldungen sind wir dankbar, denn die Tabellen wurden direkt aus unserer Oracle-Datenbank in dieses Winword-Dokument importiert. Für die Naturschutzfachstellen beider Länder haben wir die Spalte HB eingefügt, in der die Habitatsbindung entweder mit stenotop (2020 Arten mit besonderer Bindung an ausgewählte Biotopytypen) oder eurytop (1934 Arten in vielen Biotoptypen vorkommend) angegeben wurde. Somit ist belegt, dass sich Käfer besonders gut als Indikatoren für Biotope eignen. Vielleicht werden wir doch irgendwann einmal gefragt, wo die stenotopen Arten vorkommen! Gerade weil eine Klassifizierung der Käferarten in stenotop/eurytop in Fachkreisen umstritten ist, wurde von uns der KOCH in die Datenbank eingearbeitet, um damit eine Diskussion auszulösen bzw. weiterzuführen. Die Spalte M dokumentiert die Arbeiten MÜLLERs aus 1912 bzw. 1926, inklusive WASMANN (1912). Arten, die MÜLLER (1912, 1926) für das Fürstentum nachgewiesen hat, sind unter Liechtenstein mit einem x gekennzeichnet. Ein Rufezeichen (!) in den Spalten Vorarlberg bzw. Liechtenstein signalisiert eine Neumeldung. Der Punkt ( ) zeigt an, dass die Art ab 1980 in Vorarlberg nachgewiesen wurde, aber bisher in keiner der aufgeführten Publikationen - ausgenommen MÜLLER (1912, 1926) und WASMANN (1912) - erwähnt wurde. (x) = MÜLLER (1912), als unrichtig zu streichen MÜLLER (1926). Es muss hier auch festgestellt werden, dass einige Arten nur mit einem einzigen Fund eines einzigen Exemplars in diese Inventur aufgenommen wurden, obwohl ein solcher Vorgang von einem Berater der 'Vorarlberger Naturschau' als unwissenschaftlich erkannt wurde. Allen unseren - derzeit über siebzig - Mitarbeitern im In- und Ausland sei an dieser Stelle herzlich gedankt und Ihnen sei dieses Heft gewidmet. Andreas Kapp & Clemens M. Brandstetter Rankweil/Bürs, 20. März 1998.

3 Cicindelidae CICINDELA SILVICOLA DEJEAN eu x 27, 44, 52, 55 27, CICINDELA GALLICA BRULLE st x CICINDELA HYBRIDA LINNAEUS eu x 27, CICINDELA CAMPESTRIS LINNAEUS eu x 27, 31, 45, 53, 55 27, CICINDELA GERMANICA LINNAEUS eu x CICINDELA ARENARIA FUESSLIN st x Carabidae CALOSOMA INQUISITOR (LINNAEUS) eu x CALOSOMA SYCOPHANTA (LINNAEUS) eu x CARABUS CORIACEUS LINNAEUS eu x 27, 20, 31, 55, 62 21, 27, CARABUS IRREGULARIS FABRICIUS st x 27, 20, CARABUS DEPRESSUS BONELLI eu x 2, CARABUS FABRICII PANZER eu x CARABUS VIOLACEUS LINNAEUS eu x 2, 20, 27, CARABUS INTRICATUS LINNAEUS eu x CARABUS AURONITENS FABRICIUS eu x 20, 27, 45, 55 27, CARABUS PROBLEMATICUS HERBST eu x CARABUS GRANULATUS LINNAEUS eu x 27, 31, 45, 53, 55, 62 21, 27, CARABUS CANCELLATUS ILLIGER eu x 27, 31, 44, 45, 52, 55, 62 21, 27, CARABUS AURATUS LINNAEUS eu x 27, 55, 62 21, 27, CARABUS CONVEXUS FABRICIUS eu x CARABUS ARCENSIS HERBST eu x CARABUS MONILIS FABRICIUS eu x 27, 31, CARABUS SCHEIDLERI PANZER eu x 27 1, CARABUS NEMORALIS MUELLER eu x 20, 27, 55, 62 21, CARABUS HORTENSIS LINNAEUS eu x 27, CARABUS GLABRATUS PAYKULL st x 27, CARABUS CONCOLOR FABRICIUS st x CARABUS ALPESTRIS STURM st x CARABUS SILVESTRIS PANZER eu x 2, 20, CYCHRUS ANGUSTATUS HOPPE st x CYCHRUS CARABOIDES (LINNAEUS) eu x 27, 55 21, 27, CYCHRUS ATTENUATUS FABRICIUS eu x 20, 27, 55 x LEISTUS SPINIBARBIS (FABRICIUS) eu LEISTUS RUFOMARGINATUS (DUFTSCHMID) eu x LEISTUS NITIDUS (DUFTSCHMID) eu x 20, 27, 59 x LEISTUS FERRUGINEUS (LINNAEUS) eu x 27, 31, LEISTUS PICEUS FROELICH eu x 27! NEBRIA LIVIDA (LINNAEUS) eu x 27, NEBRIA PICICORNIS (FABRICIUS) st x 27, NEBRIA JOCKISCHI STURM st x 27, 55 1 Wird von AISTLEITNER & AISTLEITNER (1995) angezweifelt, ohne allerdings die Determination zu überprüfen. 1

4 NEBRIA RUFESCENS (STROEM) eu x 27 x NEBRIA BREVICOLLIS (FABRICIUS) eu x 27, 31, 38, 55, NEBRIA GERMARI HEER st x 2, NEBRIA BREMII GERMAR st x 2, NEBRIA HELLWIGI (PANZER) st NEBRIA CASTANEA (BONELLI) eu x 3, 20, NOTIOPHILUS AESTUANS MOTSCHULSKY eu x NOTIOPHILUS AQUATICUS (LINNAEUS) eu x NOTIOPHILUS PALUSTRIS (DUFTSCHMID) eu x 27, 31, 55, NOTIOPHILUS GERMINYI FAUVEL eu NOTIOPHILUS RUFIPES CURTIS eu x NOTIOPHILUS BIGUTTATUS (FABRICIUS) eu x 20, 27, 45, NOTIOPHILUS QUADRIPUNCTATUS DEJEAN 66! OMOPHRON LIMBATUM (FABRICIUS) st x ELAPHRUS CUPREUS DUFTSCHMID eu x ELAPHRUS RIPARIUS (LINNAEUS) eu x ELAPHRUS AUREUS MUELLER st x ELAPHRUS ULLRICHI REDTENBACHER st x 2, LORICERA PILICORNIS (FABRICIUS) eu x 20, 27, 55, 62 21, CLIVINA FOSSOR (LINNAEUS) eu x 27, 31, 38, 46, 53, 62 21, CLIVINA COLLARIS (HERBST) eu x 27, 31, 38, DYSCHIRIUS NITIDUS (DEJEAN) eu x DYSCHIRIUS POLITUS (DEJEAN) eu x DYSCHIRIUS AENEUS (DEJEAN) eu x 27, 31, DYSCHIRIUS INTERMEDIUS PUTZEYS eu 27, DYSCHIRIUS LAEVIUSCULUS PUTZEYS eu x DYSCHIRIUS LAFERTEI PUTZEYS eu x DYSCHIRIUS SIMILIS PETRI st 27, 31! DYSCHIRIUS GLOBOSUS (HERBST) eu x 27, 31, 46, BROSCUS CEPHALOTES (LINNAEUS) eu x MISCODERA ARCTICA (PAYKULL) eu PERILEPTUS AREOLATUS (CREUTZER) st x THALASSOPHILUS LONGICORNIS (STURM) st x 27, 44, 51, TRECHUS SECALIS (PAYKULL) eu x 27, 31, TRECHUS RUBENS (FABRICIUS) eu x TRECHUS QUADRISTRIATUS (SCHRANK) eu x 27, TRECHUS OBTUSUS ERICHSON eu x 2, 20, 27, 31, TRECHUS GLACIALIS HEER st x 2, TRECHUS TRISTIS (DUFTSCHMID) eu x LASIOTRECHUS DISCUS (FABRICIUS) eu x 27, TRECHOBLEMUS MICROS (HERBST) eu 27, 31, 44, 51, 52, TACHYS BISTRIATUS (DUFTSCHMID) st x 27, 31, 44, TACHYS MICROS (F.D.WALDHEIM) eu x TACHYS PARVULUS (DEJEAN) eu 27, TACHYS SEXSTRIATUS (DUFTSCHMID) eu x 27, 31, 44, TACHYS QUADRISIGNATUS (DUFTSCHMID) eu x 27, 31! TACHYS BISULCATUS (NICOLAI) eu! TACHYTA NANA (GYLLENHAL) eu x 27 2

5 BEMBIDION FORAMINOSUM STURM st x BEMBIDION LITORALE (OLIVIER) eu x BEMBIDION LATICOLLE (DUFTSCHMID) st x BEMBIDION PYGMAEUM (FABRICIUS) st x 27, 31, BEMBIDION LAMPROS (HERBST) eu x 27, 45, BEMBIDION PROPERANS (STEPHENS) eu x 27, 31, 38, 45, BEMBIDION PUNCTULATUM DRAPIEZ st x BEMBIDION BIPUNCTATUM (LINNAEUS) eu x BEMBIDION DENTELLUM (THUNBERG) eu x 27, 31! BEMBIDION STARKI SCHAUM st x BEMBIDION VARIUM (OLIVIER) eu x BEMBIDION SEMIPUNCTATUM (DONOVAN) eu x 27, BEMBIDION PRASINUM (DUFTSCHMID) eu x BEMBIDION TIBIALE (DUFTSCHMID) st x 27, 44, BEMBIDION GENICULATUM HEER st x 27, BEMBIDION COMPLANATUM HEER st x 27! BEMBIDION LONGIPES DANIEL st x BEMBIDION VARICOLOR (FABRICIUS) st x BEMBIDION CONFORME (DEJEAN) st x 27! BEMBIDION FASCIOLATUM (DUFTSCHMID) st x 27, BEMBIDION ASCENDENS DANIEL st x BEMBIDION MONTICOLA STURM st x 27, 44, BEMBIDION FULVIPES STURM st x BEMBIDION EQUES STURM st x BEMBIDION DELETUM SERVILLE eu x 27, 38, 55, 53! BEMBIDION INCOGNITUM MUELLER st x 2, BEMBIDION STEPHENSI CROTCH eu x BEMBIDION BRUNNICORNE DEJEAN st x BEMBIDION LUNATUM (DUFTSCHMID) st x 27, BEMBIDION TERMINALE HEER st BEMBIDION BRUXELLENSE WESMAEL eu BEMBIDION TETRACOLUM SAY eu x 27, 31, 38, BEMBIDION SUBCOSTATUM MOTSCHULSKY st BEMBIDION DISTINGUENDUM DUVAL st x BEMBIDION ANDREAE (FABRICIUS) st x 2, 27, 31, 55 27, BEMBIDION FEMORATUM STURM eu x 27, 31, BEMBIDION SCAPULARE DEJEAN st x BEMBIDION TESTACEUM (DUFTSCHMID) st x 27, BEMBIDION DECORUM (ZENKER) st x 27, BEMBIDION MODESTUM (FABRICIUS) st x BEMBIDION GLACIALE HEER st x BEMBIDION ILLIGERI NETOLITZKY st x 27, 46, 53, BEMBIDION TETRAGRAMMUM CHAUDOIR BEMBIDION STOMOIDES DEJEAN st x 27! BEMBIDION MILLERIANUM HEYDEN st x BEMBIDION RUFICORNE STURM st x BEMBIDION DECORATUM (DUFTSCHMID) st x 27, BEMBIDION DODEROI GANGLBAUER st x 27 3

6 BEMBIDION INUSTUM DUVAL st x 27! BEMBIDION SCHUEPPELI DEJEAN st x 27, BEMBIDION FUMIGATUM (DUFTSCHMID) st 27, 31, BEMBIDION ASSIMILE GYLLENHAL st x 27, BEMBIDION TENELLUM ERICHSON st x BEMBIDION AZURESCENS (DALLA TORRE) st 27, BEMBIDION HUMERALE STURM st x BEMBIDION QUADRIMACULATUM (LINNAEUS) eu x 27, 31, 46, 53, BEMBIDION DORIS (PANZER) st BEMBIDION ARTICULATUM (PANZER) eu x 27, 31, BEMBIDION QUINQUESTRIATUM GYLLENHAL st x BEMBIDION BIGUTTATUM (FABRICIUS) eu x BEMBIDION MANNERHEIMI SAHLBERG eu x 27, 31, 45, 53, BEMBIDION GUTTULA (FABRICIUS) eu x 21, BEMBIDION LUNULATUM (FOURCROY) eu ASAPHIDION CARABOIDES (SCHRANK) st x 27, 31, 55 x ASAPHIDION PALLIPES (DUFTSCHMID) eu x 27, ASAPHIDION FLAVIPES (LINNAEUS) eu x 27, 38, ASAPHIDION AUSTRIACUM SCHWEIGER 27, 55, PATROBUS AUSTRALIS SAHLBERG st x 27, PATROBUS ASSIMILIS CHAUDOIR st 2! PATROBUS ATRORUFUS (STROEM) eu x 20, 27, 31, PATROBUS STYRIACUS CHAUDOIR st 27 2, PERIGONA NIGRICEPS (DEJEAN) eu ANISODACTYLUS BINOTATUS (FABRICIUS) eu x 27, 31, 38, 55, 62 21, ANISODACTYLUS NEMORIVAGUS (DUFTSCHMID) eu x DIACHROMUS GERMANUS (LINNAEUS) eu x TRICHOTICHNUS LAEVICOLLIS (DUFTSCHMID) eu x 20, 27, TRICHOTICHNUS NITENS (HEER) eu x 20, 27, 45, PAROPHONUS MACULICORNIS (DUFTSCHMID) st HARPALUS SABULICOLA (PANZER) eu x HARPALUS MONTICOLA DEJEAN eu x HARPALUS DIFFINIS DEJEAN eu x HARPALUS NITIDULUS (STEPHENS) eu x HARPALUS RUPICOLA STURM eu x HARPALUS RUFIBARBIS (FABRICIUS) eu x 27, HARPALUS SCHAUBERGERIANUS PUEL eu HARPALUS CORDATUS (DUFTSCHMID) x HARPALUS AZUREUS (FABRICIUS) eu x HARPALUS PUNCTICOLLIS (PAYKULL) eu x HARPALUS MELLETI HEER eu 57, HARPALUS PUNCTICEPS (STEPHENS) eu 27, 31 21, HARPALUS RUFIPES (DEGEER) eu x 27, 31, 55, 62 27, HARPALUS GRISEUS (PANZER) eu x 27, 31, 38 21, HARPALUS AFFINIS (SCHRANK) eu x 27, 31, Nach Überprüfung durch Trautner & Marggi (mündl. Mitt. Marggi) P. atrorufus zuzuordnen. 4

7 HARPALUS DISTINGUENDUS (DUFTSCHMID) eu x 27, HARPALUS SMARAGDINUS (DUFTSCHMID) eu x HARPALUS DIMIDIATUS (ROSSI) eu x HARPALUS ATRATUS LATREILLE eu x HARPALUS SOLITARIS DEJEAN eu x 2, HARPALUS TENEBROSUS DEJEAN eu HARPALUS PROGREDIENS SCHAUBERGER eu HARPALUS LATUS (LINNAEUS) eu x 20, 27, 31, 55, 62 21, HARPALUS LUTEICORNIS (DUFTSCHMID) eu x HARPALUS QUADRIPUNCTATUS DEJEAN eu x 20, 27! HARPALUS RUBRIPES (DUFTSCHMID) eu x 27, 31, HARPALUS SULPHURIPES GERMAR eu x HARPALUS HONESTUS (DUFTSCHMID) eu x HARPALUS TARDUS (PANZER) eu x HARPALUS ANXIUS (DUFTSCHMID) eu x STENOLOPHUS TEUTONUS (SCHRANK) eu x 27, 31, STENOLOPHUS MIXTUS (HERBST) eu 27, 31, 44, 52 21, BRADYCELLUS RUFICOLLIS (STEPHENS) eu x BRADYCELLUS VERBASCI (DUFTSCHMID) eu x BRADYCELLUS HARPALINUS (SERVILLE) eu x BRADYCELLUS CSIKII LACZO eu BRADYCELLUS CAUCASICUS CHAUDOIR st 27, ACUPALPUS FLAVICOLLIS (STURM) eu x 27, 31, 46, ACUPALPUS MERIDIANUS (LINNAEUS) eu x 27 x ACUPALPUS PARVULUS (STURM) eu ACUPALPUS MACULATUS (SCHAUM) st STOMIS PUMICATUS (PANZER) eu x POECILUS PUNCTULATUS (SCHALLER) eu x POECILUS KOYI GERMAR st x POECILUS LEPIDUS (LESKE) eu x POECILUS CUPREUS (LINNAEUS) eu x 27, 31, 46, 53, 55, 62 21, 27, POECILUS VERSICOLOR (STURM) eu x 27, 31, 55, 62 21, 27, POECILUS CURSORIUS (DEJEAN) x PTEROSTICHUS UNCTULATUS (DUFTSCHMID) eu x 20, PTEROSTICHUS PUMILIO (DEJEAN) eu x 2, 20, 27, PTEROSTICHUS STRENUUS (PANZER) eu x 27, 31, 53, 55, 62 21, PTEROSTICHUS DILIGENS (STURM) st x 27, 31, 53, 62 21, PTEROSTICHUS OVOIDEUS (STURM) eu x 27, 53, PTEROSTICHUS VERNALIS (PANZER) eu x 27, 31, 38, 46, 53, 55, PTEROSTICHUS NIGRITA (PAYKULL) eu x 20, 27, 55, PTEROSTICHUS RHAETICUS HEER eu 27, 53, PTEROSTICHUS ANTHRACINUS (ILLIGER) eu x 27, 31, 38, PTEROSTICHUS GRACILIS (DEJEAN) eu x 27, 31, PTEROSTICHUS MINOR (GYLLENHAL) eu x 27, 31, PTEROSTICHUS OBLONGOPUNCTATUS (FABRICIUS) eu x 20, 27, 31, 45, 55, 62 x PTEROSTICHUS NIGER (SCHALLER) eu x 20, 27, 31, 55, PTEROSTICHUS MELANARIUS (ILLIGER) eu x 20, 27, 31, 38, 45, 55, PTEROSTICHUS MADIDUS (FABRICIUS) eu x 27, 55 5

8 PTEROSTICHUS AETHIOPS (PANZER) eu x 20, PTEROSTICHUS BURMEISTERI HEER x x 20, 27, 45, 55, 59 27, PTEROSTICHUS JURINEI (PANZER) eu x 20, 2, 27, PTEROSTICHUS MORIO (DUFTSCHMID) eu x PTEROSTICHUS MULTIPUNCTATUS (DEJEAN) eu x 2, 20, 27, PTEROSTICHUS FASCIATOPUNCTATUS (CREUTZER) eu x PTEROSTICHUS PANZERI (PANZER) eu x 2, MOLOPS PICEUS (PANZER) st x 20, 27, 55, ABAX PARALLELEPIPEDUS (PILLER & MITT.) eu x 20, 27, 31, 45, 55, 59, 62 27, ABAX EXARATUS DEJEAN st x ABAX PARALLELUS (DUFTSCHMID) st x 20, 27, 55, ABAX OVALIS (DUFTSCHMID) st x 20, 27, 45, 55, SYNUCHUS VIVALIS (ILLIGER) eu x 27, CALATHUS FUSCIPES (GOEZE) eu x 20, 27, 38, CALATHUS ERRATUS (SAHLBERG) eu x CALATHUS AMBIGUUS (PAYKULL) st x CALATHUS MICROPTERUS (DUFTSCHMID) eu x 20, 27 x CALATHUS MELANOCEPHALUS (LINNAEUS) eu x 27, CALATHUS OCHROPTERUS (DUFTSCHMID) st! SPHODRUS LEUCOPHTHALMUS (LINNAEUS) st x OLISTOPHUS ROTUNDATUS (PAYKULL) eu x AGONUM IMPRESSUM (PANZER) eu x x AGONUM SEXPUNCTATUM (LINNAEUS) eu x 27, 31, 53, 55, 62 21, AGONUM VIRIDICUPREUM (GOEZE) st x AGONUM GRACILIPES (DUFTSCHMID) eu x AGONUM MARGINATUM (LINNAEUS) eu x AGONUM MUELLERI (HERBST) eu x 27, 31, 38, 40, 46, 53, 55, AGONUM VIDUUM (PANZER) eu x 27, 62 21, AGONUM MOESTUM (DUFTSCHMID) eu x 27, 31, 53, 62 21, 27, AGONUM LUGENS (DUFTSCHMID) st x 27, 31, AGONUM LIVENS (GYLLENHAL) eu 27, AGONUM MICANS (NICOLAI) eu x AGONUM PICEUM (LINNAEUS) eu x 27, AGONUM GRACILE (GYLLENHAL) eu x 27, 44, AGONUM FULIGINOSUM (PANZER) eu x 27, 31, 53 21, AGONUM PELIDNUM (PAYKULL) st x 27, 31, PLATYNUS ASSIMILIS (PAYKULL) eu x 20, 27, 31, 38, 55, 59, PLATYNUS LONGIVENTRIS (MANNERHEIM) st x 27, PLATYNUS ALBIPES (FABRICIUS) eu x 27, 31, 38, PLATYNUS OBSCURUS (HERBST) eu x 27, PLATYNUS DORSALIS (PONTOPPIDAN) st x 27, 45, 53, 55, 62! ZABRUS TENEBRIOIDES (GOEZE) st x AMARA PLEBEJA (GYLLENHAL) eu x AMARA SIMILATA (GYLLENHAL) eu x 27, 31, 38, AMARA OVATA (FABRICIUS) eu x 27, 28, 31, AMARA MONTIVAGA STURM eu x 27, 55 27, AMARA NITIDA STURM eu x AMARA CONVEXIOR STEPHENS eu x 27,

9 AMARA COMMUNIS (PANZER) eu x 27, 31, 62 21, AMARA SCHIMPERI WENCKER st x 27, 66 x AMARA CURTA DEJEAN eu x AMARA LUNICOLLIS SCHIOEDTE eu x 27, 31, AMARA AENEA (DEGEER) eu x 27, 31, 55, 62 21, AMARA EYRINOTA (PANZER) eu x AMARA FAMILIARIS (DUFTSCHMID) eu x 27, 31, AMARA LUCIDA (DUFTSCHMID) eu AMARA ERRATICA (DUFTSCHMID) eu x 20, AMARA QUENSELI (SCHOENHERR) st x 2, AMARA BIFRONS (GYLLENHAL) eu x AMARA PRAETERMISSA (SAHLBERG) eu x 2, 27! AMARA FULVA (MUELLER) eu x AMARA CONSULARIS (DUFTSCHMID) eu x AMARA AULICA (PANZER) eu x 20, 27, 31, 55, 62 21, AMARA HELLERI GREDLER st AMARA EQUESTRIS (DUFTSCHMID) eu x CHLAENIUS TRISTIS (SCHALLER) eu x CHLAENIUS NIGRICORNIS (FABRICIUS) eu x 27, 31, 38, 44, 52 21, CHLAENIUS NITIDULUS (SCHRANK) eu x CHLAENIUS TIBIALIS (DEJEAN) st x 27, 31, CHLAENIUS VESTITUS (PAYKULL) eu x CALLISTUS LUNATUS (FABRICIUS) eu x 27, OODES HELOPIOIDES (FABRICIUS) st x 27, 31, 62 21, LICINUS HOFFMANNSEGGI (PANZER) eu LICINUS DEPRESSUS (PAYKULL) eu x x BADISTER BULLATUS (SCHRANK) eu x BADISTER LACERTOSUS STURM eu x 27, 31, 53 21, BADISTER MERIDIONALIS PUEL eu BADISTER SODALIS (DUFTSCHMID) eu x 27, 31, 45, 53, 62 21, BADISTER DILATATUS CHAUDOIR st BADISTER PELTATUS (PANZER) st x 27, BADISTER ANOMALUS (PERRIS) st 31, PANAGAEUS CRUXMAJOR (LINNAEUS) st x 27, 31, 44, PANAGAEUS BIPUSTULATUS (FABRICIUS) st x 44, ODACANTHA MELANURA (LINNAEUS) st 27, 31, LEBIA CHLOROCEPHALA (HOFFMANN) eu x LEBIA CYANOCEPHALA (LINNAEUS) st x LEBIA CRUXMINOR (LINNAEUS) eu x 27, LEBIA MARGINATA (FOURCROY) eu x DEMETRIAS ATRICAPILLUS (LINNAEUS) eu DEMETRIAS IMPERIALIS (GERMAR) st 27, CYMINDIS HUMERALIS (FOURCROY) eu x CYMINDIS AXILLARIS (FABRICIUS) eu x CYMINDIS VAPORARIORUM (LINNAEUS) st x DROMIUS LONGICEPS DEJEAN st 27, 31! DROMIUS LINEARIS (OLIVIER) eu x 27, 31, 44, 46, 52, 53, DROMIUS AGILIS (FABRICIUS) eu x 20, 27 7

10 DROMIUS ANGUSTUS BRULLE eu DROMIUS FENESTRATUS (FABRICIUS) eu x 20, DROMIUS QUADRIMACULATUS (LINNAEUS) eu x 27, 31, 45, 53, DROMIUS SPILOTUS (ILLIGER) eu x DROMIUS NOTATUS STEPHENS eu x 27, SYNTOMUS TRUNCATELLUS (LINNAEUS) eu LIONYCHUS QUADRILLUM (DUFTSCHMID) st x MICROLESTES MINUTULUS (GOEZE) eu x 27, BRACHINUS CREPITANS (LINNAEUS) st x Haliplidae BRYCHIUS ELEVATUS (PANZER) st x 10, PELTODYTES CAESUS (DUFTSCHMID) eu 10 x HALIPLUS OBLIQUUS (FABRICIUS) st x 10, 49 x HALIPLUS CONFINIS STEPHENS st x 10, 31, HALIPLUS LINEATOCOLLIS (MARSHAM) eu x 10, 49, HALIPLUS RUFICOLLIS (DEGEER) eu x 10, 31, HALIPLUS HEYDENI WEHNCKE eu x 31, 44, 45, 49, 52, 53, 62 21, HALIPLUS FLUVIATILIS AUBE st x 10, 49, HALIPLUS WEHNCKEI GERHARDT eu 31, 49, HALIPLUS FULVICOLLIS ERICHSON eu x HALIPLUS LAMINATUS (SCHALLER) eu x 10, 49, HALIPLUS MUCRONATUS STEPHENS st x HALIPLUS FLAVICOLLIS STURM eu x 10, 31, 45, 49, HALIPLUS FULVUS (FABRICIUS) st x 10, HALIPLUS VARIEGATUS STURM st x 10 Noteridae NOTERUS CLAVICORNIS (DEGEER) eu x 10, 25, 31, 46, 49, 53 25, NOTERUS CRASSICORNIS (MUELLER) eu x 10, 25, 31, 49 Dytiscidae HYPHYDRUS OVATUS (LINNAEUS) eu x 25, 31, HYDROGLYPHUS PUSILLUS (FABRICIUS) eu x 10, 25, 31, 46, 49, 53, 62 21, BIDESSUS MINUTISSIMUS (GERMAR) st 24, 25, 31, 49, BIDESSUS DELICATULUS (SCHAUM) st 25, BIDESSUS UNISTRIATUS (SCHRANK) st x 10, 25, 31, BIDESSUS GROSSEPUNCTATUS VORBRINGER st 25, 31, 49, COELAMBUS IMPRESSOPUNCTATUS (SCHALLER) eu x 10, 25, 31, 49, 53, COELAMBUS PARALLELOGRAMMUS (AHRENS) st COELAMBUS CONFLUENS (FABRICIUS) eu 25, HYGROTUS VERSICOLOR (SCHALLER) eu x 10, 25, 31, HYGROTUS INAEQUALIS (FABRICIUS) eu x 10, 25, 31, 49 21, 25, 49 8

11 HYGROTUS QUINQUELINEATUS (ZETTERSTEDT) st 7, 14, 17, 24, 25, 31, HYGROTUS DECORATUS (GYLLENHAL) eu 25, 31, 49, HYDROPORUS SCALESIANUS STEPHENS st 31, HYDROPORUS ANGUSTATUS STURM eu x 10, 25, 31, 49, 52, 53, 62 21, HYDROPORUS UMBROSUS (GYLLENHAL) st 31, HYDROPORUS TRISTIS (PAYKULL) st x 10, 25, 31, 49, 53, 62 x HYDROPORUS NOTATUS STURM st HYDROPORUS PICEUS STEPHENS st HYDROPORUS PALUSTRIS (LINNAEUS) eu x 10, 25, 31, 45, 49, 53, 62 25, HYDROPORUS INCOGNITUS SHARP st 10, 25, 31, 49, 52, 53, HYDROPORUS STRIOLA (GYLLENHAL) st 31, HYDROPORUS ERYTHROCEPHALUS (LINNAEUS) eu x 10, 25, 31, 49, 53, 62! HYDROPORUS MARGINATUS (DUFTSCHMID) st x 10, 25, HYDROPORUS RUFIFRONS (DUFTSCHMID) eu x 10, 25, 31, 49 x HYDROPORUS PLANUS (FABRICIUS) eu x 10, 25, HYDROPORUS PUBESCENS (GYLLENHAL) eu HYDROPORUS DISCRETUS FAIRMAIRE st x 10, 25, 49, 53, 62 25, HYDROPORUS FOVEOLATUS HEER st x 10, 25, HYDROPORUS NIVALIS HEER st x 2, 10, 25, HYDROPORUS NIGRITA (FABRICIUS) eu x 10, 25, 49, HYDROPORUS MEMNONIUS NICOLAI st x 10, 25, 31, 49, 53, 62 x HYDROPORUS FERRUGINEUS STEPHENS st x 10, 25, HYDROPORUS MELANARIUS STURM st x 10, 25, 31, 49, 52, 53, 62 x HYDROPORUS LONGICORNIS SHARP st 49, HYDROPORUS NEGLECTUS SCHAUM st x 10, 25, 31, 49, HYDROPORUS KRAATZI SCHAUM st 25, GRAPTODYTES GRANULARIS (LINNAEUS) eu x 10, 25, 31, 49, GRAPTODYTES BILINEATUS (STURM) eu x GRAPTODYTES PICTUS (FABRICIUS) eu x 10, 25, 46, 49, 52, 53, 62 25, PORHYDRUS LINEATUS (FABRICIUS) eu x 10, 25, 31, DERONECTES AUBEI (MULSANT) st DERONECTES PLATYNOTUS (GERMAR) st 25, 30, DERONECTES LATUS (STEPHENS) st STICTOTARSUS DUODECIMPUSTULATUS (FABRICIUS) eu x 10, 25, 49, NEBRIOPORUS GRISEOSTRIATUS (DEGEER) st x 2, 8, 10, 25, NEBRIOPORUS DEPRESSUS (FABRICIUS) st x 10, 25, 44, 49, NEBRIOPORUS ASSIMILIS (PAYKULL) st x 10, 25, OREODYTES DAVISII (CURTIS) st x 10, 25, OREODYTES SEPTENTRIONALIS (GYLLENHAL) st x 10, 25, OREODYTES SANMARKII (SAHLBERG) st x 10, 25, 44, 49, SCARODYTES HALENSIS (FABRICIUS) eu LACCOPHILUS VARIEGATUS (GERMAR) st x 10, 25, 31, LACCOPHILUS MINUTUS (LINNAEUS) eu x 10, 25, 46, 49, 53, LACCOPHILUS HYALINUS (DEGEER) eu 10, 25, 31, 46, 49, COPELATUS HAEMORRHOIDALIS (FABRICIUS) eu x 10, 25, 31, 49, 53, 62 21, PLATAMBUS MACULATUS (LINNAEUS) st x 10, 25, 49, 62 25, AGABUS CHALCONATUS (PANZER) st x 10, AGABUS NEGLECTUS ERICHSON st 25, 49 9

12 AGABUS BIGUTTATUS (OLIVIER) st x 10, 25, AGABUS NITIDUS (FABRICIUS) st x 10, 25, 49, 62! AGABUS GUTTATUS (PAYKULL) st x 10, 25, 49, AGABUS MELANARIUS AUBE st x 10, 25, AGABUS BIPUSTULATUS (LINNAEUS) eu x 10, 25, 31, 49, 53, AGABUS SOLIERI AUBE st x 10, 25, AGABUS STURMII (GYLLENHAL) eu x 10, 25, 31, 45, 49, AGABUS ULIGINOSUS (LINNAEUS) eu AGABUS PALUDOSUS (FABRICIUS) eu x 10, 25, 49, 52, AGABUS NEBULOSUS (FORSTER) eu x AGABUS AFFINIS (PAYKULL) st 16, 25, 31, 49, 52, AGABUS CONGENER (THUNBERG) st x 10, 25, AGABUS DIDYMUS (OLIVIER) st 25, 49, 53, AGABUS UNDULATUS (SCHRANK) eu ILYBIUS FENESTRATUS (FABRICIUS) eu x 10, 25, 49, ILYBIUS ATER (DEGEER) st x 10, 25, 31, 49, ILYBIUS FULIGINOSUS (FABRICIUS) eu x 10, 25, 49, 62 21, 25, ILYBIUS CRASSUS THOMSON st! ILYBIUS QUADRIGUTTATUS (LACORDAIRE) eu x 10, ILYBIUS GUTTIGER (GYLLENHAL) st x 10, 31, 49, ILYBIUS AENESCENS THOMSON st NARTUS GRAPEI (GYLLENHAL) st x 10, 25, 31, 49, 52, RHANTUS SUTURALIS (M'LEAY) eu x 10, 25, 31, 49, 53, 62 21, RHANTUS FRONTALIS (MARSHAM) eu RHANTUS BISTRIATUS (BERGSTRAESSER) eu x RHANTUS CONSPUTUS (STURM) st RHANTUS EXSOLETUS (FORSTER) eu x 10, RHANTUS LATITANS SHARP st x 10, 25, 31, COLYMBETES FUSCUS (LINNAEUS) eu x HYDATICUS TRANSVERSALIS (PONTOPPIDAN) eu x 10, 25, 31, HYDATICUS SEMINIGER (DEGEER) eu x 10, 25, 31, 49, HYDATICUS GRAMMICUS (GERMAR) st GRAPHODERUS BILINEATUS (DEGEER) eu x 10, GRAPHODERUS ZONATUS (HOPPE) st x GRAPHODERUS CINEREUS (LINNAEUS) eu x 10, 25, 31, ACILIUS SULCATUS (LINNAEUS) eu x 10, 25, 49, DYTISCUS LATISSIMUS LINNAEUS st x DYTISCUS SEMISULCATUS MUELLER eu x DYTISCUS MARGINALIS LINNAEUS eu x 10, 25, 31, 45, 49, 52, 53, 63 21, 25, CYBISTER LATERALIMARGINALIS (DEGEER) eu x 10 Gyrinidae GYRINUS MINUTUS FABRICIUS eu x GYRINUS MARINUS GYLLENHAL eu x 10, GYRINUS COLYMBUS ERICHSON eu x 10, GYRINUS SUBSTRIATUS STEPHENS eu 49, 53,

13 GYRINUS NATATOR (LINNAEUS) st x GYRINUS SUFFRIANI SCRIBA st x 10, GYRINUS PAYKULLI OCHS st 49, ORECTOCHILUS VILLOSUS MUELLER st x 10, 49, Microsporidae MICROSPORUS OBSIDIANUS KOLENATI eu x Hydrochidae HYDROCHUS ELONGATUS (SCHALLER) eu x HYDROCHUS CARINATUS GERMAR eu x HYDROCHUS BREVIS (HERBST) st x HYDROCHUS MEGAPHALLUS BERGE HYDROCHUS IGNICOLLIS MOTSCHULSKY 62 Hydraenidae HYDRAENA PALUSTRIS ERICHSON eu x 31, HYDRAENA BRITTENI JOY st! HYDRAENA RIPARIA KUGELANN eu x HYDRAENA MELAS DALLA TORRE st 31, 53, 62! HYDRAENA NIGRITA GERMAR st! HYDRAENA PYGMAEA WATERHOUSE st x HYDRAENA LAPIDICOLA KIESENWETTER st x! HYDRAENA GRACILIS GERMAR st x HYDRAENA SAGA D'ORCHYMONT st! C HYDRAENA SAGA ALPICOLA PRETNER st! HYDRAENA TRUNCATA REY st HYDRAENA POLITA KIESENWETTER st x HYDRAENA DENTIPES GERMAR st! HYDRAENA MINUTISSIMA STEPHENS eu x OCHTHEBIUS GRANULATUS MULSANT st x OCHTHEBIUS EXSCULPTUS GERMAR st x OCHTHEBIUS GIBBOSUS GERMAR st x 44, OCHTHEBIUS BICOLON GERMAR eu 44, 52, 53, OCHTHEBIUS CRENULATUS MULSANT & REY 50, OCHTHEBIUS MINIMUS (FABRICIUS) eu! OCHTHEBIUS METALLESCENS ROSENHAUER st! OCHTHEBIUS SIDANUS ORCHYMONT st x OCHTHEBIUS NOBILIS VILLA st x LIMNEBIUS TRUNCATELLUS THUNBERG st x 53, LIMNEBIUS PAPPOSUS MULSANT eu x LIMNEBIUS NITIDUS (MARSHAM) st x LIMNEBIUS ALUTA BEDEL st x 34 11

14 LIMNEBIUS ATOMUS (DUFTSCHMID) st x 31, 34 Spercheidae SPERCHEUS EMARGINATUS (SCHALLER) st x 31 Georissidae GEORISSUS CRENULATUS (ROSSI) x GEORISSUS LAESICOLLIS GERMAR x Hydrophilidae HELOPHORUS PORCULUS BEDEL st x HELOPHORUS NUBILUS FABRICIUS eu x HELOPHORUS SCHMIDTI VILLA st x HELOPHORUS GRANDIS ILLIGER eu! HELOPHORUS AQUATICUS (LINNAEUS) eu x 31, 53, HELOPHORUS ARVERNICUS MULSANT st HELOPHORUS NIVALIS GIRAUD st x! HELOPHORUS GLACIALIS VILLA st x! HELOPHORUS BREVIPALPIS BEDEL eu x 31, HELOPHORUS NANUS STURM st x HELOPHORUS PUMILIO ERICHSON st x 31, HELOPHORUS REDTENBACHERI KUWERT st 44, 51, 52, HELOPHORUS FLAVIPES FABRICIUS eu x HELOPHORUS OBSCURUS MULSANT st HELOPHORUS ASPERATUS REY st x 44, 52, HELOPHORUS DORSALIS (MARSHAM) st! HELOPHORUS GRANULARIS (LINNAEUS) eu x 31, 46, 53, HELOPHORUS MINUTUS FABRICIUS eu 53, HELOPHORUS GRISEUS HERBST st x COELOSTOMA ORBICULARE (FABRICIUS) st x 31, 53, 62 x SPHAERIDIUM BIPUSTULATUM FABRICIUS eu x SPHAERIDIUM SUBSTRIATUM FALDERMANN st x SPHAERIDIUM SCARABAEOIDES (LINNAEUS) eu x 31! SPHAERIDIUM LUNATUM FABRICIUS eu x 53, SPHAERIDIUM MARGINATUS FABRICIUS! CERCYON USTULATUS (PREYSSLER) eu x CERCYON OBSOLETUS GYLLENHAL eu x CERCYON IMPRESSUS (STURM) eu x 45, 62! CERCYON HAEMORRHOIDALIS (FABRICIUS) eu x 28, CERCYON ALPINUS VOGT st CERCYON MELANOCEPHALUS (LINNAEUS) eu x 45, 53! CERCYON MARINUS THOMSON st x CERCYON BIFENESTRATUS KUESTER st! 12

15 CERCYON LATERALIS (MARSHAM) eu x 31, 53, CERCYON LAMINATUS SHARP eu 31, CERCYON UNIPUNCTATUS (LINNAEUS) eu x CERCYON QUISQUILIUS (LINNAEUS) eu x CERCYON ATRICAPILLUS (MARSHAM) eu x x CERCYON TERMINATUS (MARSHAM) eu x CERCYON PYGMAEUS (ILLIGER) eu x 31, 53, CERCYON GRANARIUS ERICHSON st x CERCYON TRISTIS (ILLIGER) eu x CERCYON CONVEXIUSCULUS STEPHENS eu x CERCYON STERNALIS SHARP eu CERCYON ANALIS (PAYKULL) eu x MEGASTERNUM OBSCURUM (MARSHAM) eu x 31, 45, 53, 62! CRYPTOPLEURUM MINUTUM (FABRICIUS) eu x 31, 53, 62! CRYPTOPLEURUM CRENATUM (PANZER) st x CRYPTOPLEURUM SUBTILE SHARP eu CRENITIS PUNCTATOSTRIATA (LETZNER) st! HYDROBIUS FUSCIPES (LINNAEUS) eu x 31, 45, 53, ANACAENA GLOBULUS (PAYKULL) eu x 31, 53, ANACAENA LIMBATA (FABRICIUS) eu x 31, 46, 53, ANACAENA LUTESCENS (STEPHENS) eu 31, 45, 53, LACCOBIUS STRIATULUS (FABRICIUS) st x 31, 46, 53, LACCOBIUS SINUATUS MOTSCHULSKY eu x LACCOBIUS OBSCURATUS (ROTTENBERG) st x LACCOBIUS NEAPOLITANUS ROTTENBERG st! LACCOBIUS ATROCEPHALUS REITTER LACCOBIUS BIPUNCTATUS (FABRICIUS) eu x 46, LACCOBIUS MINUTUS (LINNAEUS) eu x 31, 46, 62 x LACCOBIUS BIGUTTATUS GERHARDT st! LACCOBIUS GRACILIS MOTSCHULSKY st x LACCOBIUS ALTERNUS MOTSCHULSKY st x 31 x HELOCHARES LIVIDUS (FORSTER) eu x x HELOCHARES OBSCURUS (MUELLER) eu x 31, 62! ENOCHRUS MELANOCEPHALUS (OLIVIER) st x ENOCHRUS OCHROPTERUS (MARSHAM) st x x ENOCHRUS QUADRIPUNCTATUS (HERBST) eu x 31, A ENOCHRUS QUADRIPUNCTATUS VAR.FUSCIPENNIS x ENOCHRUS TESTACEUS (FABRICIUS) eu x 31, ENOCHRUS AFFINIS (THUNBERG) st x ENOCHRUS COARCTATUS (GREDLER) st x 31! CYMBIODYTA MARGINELLA (FABRICIUS) st x CAETARTHRIA SEMINULUM (HERBST) eu x HYDROCHARA CARABOIDES (LINNAEUS) x 31, 53, HYDROPHILUS PICEUS (LINNAEUS) x HYDROPHILUS ATERRIMUS (ESCHSCHOLTZ) st x BEROSUS SIGNATICOLLIS (CHARPENTIER) st x 31, 35, BEROSUS LURIDUS (LINNAEUS) eu x 31, 35 13

16 Histeridae TERETRIUS PICIPES (FABRICIUS) st x PLEGADERUS VULNERATUS (PANZER) st! PLEGADERUS CAESUS (HERBST) eu ONTHOPHILUS PUNCTATUS (MUELLER) eu x ABRAEUS GRANULUM ERICHSON eu! ABRAEUS PARVULUS AUBE st ACRITUS NIGRICORNIS (HOFFMANN) eu x MYRMETES PAYKULLI (KANAAR) st x GNATHONCUS ROTUNDATUS (KUGELANN) eu x GNATHONCUS NANNETENSIS (MARSEUL) eu! GNATHONCUS BUYSSONI AUZAT eu SAPRINUS SEMISTRIATUS (SCRIBA) eu x SAPRINUS LAUTUS ERICHSON st x SAPRINUS AENEUS (FABRICIUS) eu x HYPOCACCUS RUGIFRONS (PAYKULL) eu x HYPOCACCUS METALLICUS (HERBST) eu x HYPOCACCUS RUGICEPS (DUFTSCHMID) st x DENDROPHILUS PUNCTATUS (HERBST) eu x DENDROPHILUS PYGMAEUS (LINNAEUS) eu x CARCINOPS PUMILIO (ERICHSON) eu x PAROMALUS FLAVICORNIS (HERBST) eu! PAROMALUS PARALLELEPIPEDUS (HERBST) eu x HOLOLEPTA PLANA (SULZER) st x PLATYSOMA MINOR (ROSSI) st x GRAMMOSTETHUS MARGINATUS (ERICHSON) x PARALISTER OBSCURUS (KUGELANN) eu x PARALISTER PURPURASCENS (HERBST) eu x PARALISTER NEGLECTUS (GERMAR) eu x PARALISTER VENTRALIS (MARSEUL) eu x PARALISTER CARBONARIUS (HOFFMANN) eu x HISTER QUADRIMACULATUS LINNAEUS eu x HISTER UNICOLOR LINNAEUS eu x HISTER STRIOLA SAHLBERG x HISTER QUADRINOTATUS SCRIBA eu x HISTER TERRICOLA GERMAR x HISTER MERDARIUS HOFFMANN x HISTER BRUNNEUS (FABRICIUS) x HISTER ILLIGERI DUFTSCHMID st x HISTER HELLUO TRUQUI st x HISTER FUNESTUS ERICHSON eu x HISTER BISSEXSTRIATUS FABRICIUS eu x ATHOLUS BIMACULATUS (LINNAEUS) eu x ATHOLUS DUODECIMSTRIATUS (SCHRANK) eu x ATHOLUS CORVINUS (GERMAR) eu x 14

17 HETAERIUS FERRUGINEUS (OLIVIER) eu x Sphaeritidae SPHAERITES GLABRATUS (FABRICIUS) st x Silphidae NECROPHORUS HUMATOR (GLEDITSCH) eu x! NECROPHORUS INVESTIGATOR ZETTERSTEDT eu x x NECROPHORUS FOSSOR ERICHSON eu x NECROPHORUS VESPILLOIDES HERBST eu x 29, NECROPHORUS VESPILLO (LINNAEUS) eu x NECRODES LITTORALIS (LINNAEUS) eu x THANATOPHILUS RUGOSUS (LINNAEUS) eu x THANATOPHILUS SINUATUS (FABRICIUS) eu x THANATOPHILUS DISPAR (HERBST) eu x OECEOPTOMA THORACICA (LINNAEUS) eu x 29, 45, BLITHOPHAGA OPACA (LINNAEUS) st! BLITHOPHAGA UNDATA (MUELLER) st x XYLODREPA QUADRIMACULATA (SCOPOLI) eu x SILPHA OBSCURA LINNAEUS eu x SILPHA TRISTIS ILLIGER eu x SILPHA TYROLENSIS LAICHARTING eu x! ABLATTARIA LAEVIGATA (FABRICIUS) eu x x PHOSPHUGA ATRATA (LINNAEUS) eu x 45, 53, Agyrtidae NECROPHILUS SUBTERRANEUS (DAHL) st x AGYRTES BICOLOR L.D.CASTELNAU eu 43 Leptinidae LEPTINUS TESTACEUS MUELLER eu 19 Cholevidae PTOMAPHAGUS VARIICORNIS (ROSENHAUER) eu x 45, 53, PTOMAPHAGUS SUBVILLOSUS (GOEZE) eu x PTOMAPHAGUS SERICATUS (CHAUDOIR) eu x NEMADUS COLONOIDES (KRAATZ) eu! NARGUS VELOX (SPENCE) eu x 62! NARGUS BADIUS (STURM) st x NARGUS WILKINI (SPENCE) eu 45, 62! 15

18 NARGUS ANISOTOMOIDES (SPENCE) eu x! CHOLEVA OBLONGA LATREILLE eu! CHOLEVA NIVALIS (KRAATZ) eu! CHOLEVA CISTELOIDES (FROELICH) eu x CHOLEVA GLAUCA BRITTEN eu CHOLEVA ANGUSTATA (FABRICIUS) eu! CHOLEVA STURMI BRISOUT st! CHOLEVA FAGNIEZI JEANNEL eu! CHOLEVA ELONGATA (PAYKULL) eu x DREPOSCIA RELICTA LOHSE st SCIODREPOIDES WATSONI (SPENCE) eu x 45, 53, SCIODREPOIDES FUMATUS (SPENCE) eu x 29, CATOPS SUBFUSCUS KELLNER st x CATOPS LONGULUS KELLNER st CATOPS CORACINUS KELLNER eu x CATOPS KIRBYI (SPENCE) eu! CATOPS TRISTIS (PANZER) eu x! CATOPS CHRYSOMELOIDES (PANZER) eu x CATOPS NEGLECTUS KRAATZ st! CATOPS MORIO (FABRICIUS) eu x CATOPS NIGRITA ERICHSON eu x 53, CATOPS NIGRICLAVIS GERHARDT eu x 2! CATOPS JOFFREI DEVILLE st CATOPS WESTI KROG st 45, 53, CATOPS FUSCUS (PANZER) eu! CATOPS FULIGINOSUS ERICHSON eu x 53, 62! CATOPS NIGRICANS (SPENCE) eu x 2, 53, CATOPS PICIPES (FABRICIUS) eu x! Colonidae COLON LATUM KRAATZ eu x COLON GRISEUM CZWALINA eu x COLON ANGULARE ERICHSON eu x COLON DENTIPES (SAHLBERG) st x! COLON BRUNNEUM (LATREILLE) eu x COLON APPENDICULATUM (SAHLBERG) eu x COLON CALCARATUM ERICHSON st x Leiodidae TRIARTHRON MAERKELI MAERKEL eu! HYDNOBIUS SPINIPES (GYLLENHAL) eu! HYDNOBIUS PUNCTATUS (STURM) eu x HYDNOBIUS MULTISTRIATUS (GYLLENHAL) eu x LEIODES PALLENS (STURM) st x LEIODES SUBCONVEXA DAFFNER! 16

19 LEIODES ROTUNDATA (ERICHSON) st x LEIODES TRIEPKEI (SCHMIDT) eu x LEIODES RUGOSA STEPHENS eu! LEIODES BRANDISI HOLDHAUS st LEIODES CINNAMOMEA (PANZER) eu x LEIODES OBLONGA (ERICHSON) st! LEIODES RHAETICA ERICHSON! LEIODES FLAVESCENS (SCHMIDT) eu x LEIODES POLITA (MARSHAM) eu x! LEIODES PICEA (PANZER) eu x LEIODES DUBIA (KUGELANN) eu x LEIODES RUFIPENNIS (PAYKULL)! LEIODES RUFICOLLIS SAHLBERG eu x LEIODES FERRUGINEA (FABRICIUS) eu x LEIODES GALLICA REITTER LEIODES BADIA (STURM) st x COLENIS IMMUNDA (STURM) eu x AGARICOPHAGUS CEPHALOTES SCHMIDT eu x CYRTUSA MINUTA (AHRENS) eu x CYRTUSA PAUXILLA (SCHMIDT) x CYRTUSA SUBTESTACEA (GYLLENHAL) x CYRTUSA MINUTA (AHRENS)! ANISOTOMA HUMERALIS (FABRICIUS) eu x ANISOTOMA AXILLARIS GYLLENHAL eu x ANISOTOMA CASTANEA (HERBST) eu x ANISOTOMA GLABRA (KUGELANN) st x ANISOTOMA ORBICULARIS (HERBST) eu x LIODOPRIA SERRICORNIS (GYLLENHAL) st x AMPHICYLLIS GLOBUS (FABRICIUS) eu x! AMPHICYLLIS GLOBIFORMIS (SAHLBERG) st x CYRTOPLASTUS SERIEPUNCTATUS (BRISOUT) st 43, 44, 51, AGATHIDIUM MARGINATUM STURM eu x AGATHIDIUM VARIANS (BECK) eu x 62! AGATHIDIUM MANDIBULARE STURM eu x AGATHIDIUM ROTUNDATUM (GYLLENHAL) eu x AGATHIDIUM CONFUSUM BRISOUT eu AGATHIDIUM ARCTICUM THOMSON st AGATHIDIUM NIGRIPENNE (FABRICIUS) eu x! AGATHIDIUM ATRUM (PAYKULL) eu!! AGATHIDIUM SEMINULUM (LINNAEUS) eu! AGATHIDIUM LAEVIGATUM ERICHSON eu 62! AGATHIDIUM DENTATUM MULSANT & REY eu x 45! AGATHIDIUM BOHEMICUM REITTER eu! AGATHIDIUM BADIUM ERICHSON eu! AGATHIDIUM PISANUM BRISOUT! Scydmaenidae 17

20 EUTHIA LINEARIS MULSANT eu! CEPHENNIUM THORACICUM MUELLER&KUNZE eu x NEURAPHES ELONGATULUS (MUELLER&KUNZE) eu x 45, 53, NEURAPHES ANGULATUS (MUELLER&KUNZE) eu! NEURAPHES CARINATUS (MULSANT) eu x NEURAPHES TALPARUM LOKAY eu! NEURAPHES PLICICOLLIS REITTER eu! NEURAPHES CORONATUS SAHLBERG eu NEURAPHES CAVICEPS CROISSANDEAU st NEURAPHES PARALLELUS (CHAUDOIR) st x SCYDMORAPHES SPARSHALLI (DENNY) eu!! SCYDMORAPHES HELVOLUS (SCHAUM) eu! STENICHNUS SCUTELLARIS (MUELLER&KUNZE) eu x 53, 62! STENICHNUS PUSILLUS (MUELLER&KUNZE) st x STENICHNUS GODARTI LATREILLE st! STENICHNUS COLLARIS (MUELLER&KUNZE) eu x 45, 53, STENICHNUS BICOLOR (DENNY) eu! MICROSCYDMUS NANUS (SCHAUM) eu!! EUCONNUS PUBICOLLIS (MUELLER&KUNZE) eu x 53, EUCONNUS CLAVIGER (MUELLER&KUNZE) eu! EUCONNUS WETTERHALLI (GYLLENHAL) eu x 46, 62! EUCONNUS CARINTHIACUS GANGLBAUER st! EUCONNUS DENTICORNIS (MUELLER&KUNZE) st EUCONNUS HIRTICOLLIS (ILLIGER) st x SCYDMAENUS TARSATUS MUELLER&KUNZE eu x 46! SCYDMAENUS RUFUS MUELLER&KUNZE eu! SCYDMAENUS PERRISI REITTER st! Ptiliidae NOSSIDIUM PILOSELLUM MARSHAM st PTENIDIUM GRESSNERI ERICHSON eu! PTENIDIUM LAEVIGATUM ERICHSON st! PTENIDIUM TURGIDUM THOMSON st! PTENIDIUM INTERMEDIUM WANKOWICZ st x 53! PTENIDIUM BRENSKEI FLACH st PTENIDIUM FUSCICORNE ERICHSON st x PTENIDIUM FORMICETORUM KRAATZ eu x PTENIDIUM PUSILLUM (GYLLENHAL) eu x PTENIDIUM MATTHEWSI FLACH x PTENIDIUM NITIDUM (HEER) x ACTIDIUM BOUDIERI (ALLIBERT) st x PTILIUM MYRMECOPHILUM (ALLIBERT) eu! PTILIUM MODESTUM WANKOWICZ eu! EURYPTILIUM GILLMEISTERI FLACH st EURYPTILIUM SAXONICUM (GILLMEISTER) eu! 18

21 PTILIOLA KUNZEI (HEER) eu x PTILIOLA BREVICOLLE (MATTHEWS) st! PTILIOLUM SPENCEI (ALLIBERT) eu x PTILIOLUM FUSCUM (ERICHSON) eu! PTILIOLUM MARGINATUM (AUBE) st PTILIOLUM SAHLBERGI FLACH st PTINELLA BRITANNICA MATTHEWS st! PTINELLA LIMBATA (HEER) eu! PTINELLA APTERA (GUERIN-MEN.) eu! PTERYX SUTURALIS (HEER) eu 62! NEPHANES TITAN (NEWMAN) eu!! SMICRUS FILICORNIS (FAIRMAIRE&LAB.) st x BAEOCRARA VARIOLOSA (MULSANT & REY eu x BAEOCRARA JAPONICA (MATTHEWS) ACROTRICHIS GRANDICOLLIS (MANNERHEIM) eu x 53 x ACROTRICHIS MONTANDONI (ALLIBERT) eu x! ACROTRICHIS THORACICA (WALTL) eu x! ACROTRICHIS SERICANS (HEER) eu x ACROTRICHIS DISPAR (MATTHEWS) eu! ACROTRICHIS BREVIPENNIS (ERICHSON) eu x ACROTRICHIS PUMILA (ERICHSON) eu ACROTRICHIS SILVATICA ROSSKOTHEN eu 2! ACROTRICHIS PARVA ROSSKOTHEN eu 2! ACROTRICHIS VOLANS (MOTSCHULSKY) st 45! ACROTRICHIS ARNOLDI ROSSKOTHEN eu ACROTRICHIS INTERMEDIA (GILLMEISTER) eu x 45, 53, 62! ACROTRICHIS ATOMARIA (DEGEER) eu x 53! ACROTRICHIS SITKAENSIS (MOTSCHULSKY) eu!! ACROTRICHIS ROSSKOTHENI SUNDT st ACROTRICHIS FASCICULARIS (HERBST) eu x 53, 62! ACROTRICHIS RUGULOSA ROSSKOTHEN eu Dasyceridae DASYCERUS SULCATUS BRONGNIART eu x 45, 62 Scaphidiidae SCAPHIDIUM QUADRIMACULATUM OLIVIER eu x 45, 62! SCAPHISOMA AGARICINUM (LINNAEUS) eu x 45, 53, SCAPHISOMA BOLETI (PANZER) st x SCAPHISOMA ASSIMILE ERICHSON st x SCAPHISOMA BOREALE LUNDBLAD 21 Micropeplidae 19

22 MICROPEPLUS TESSERULA CURTIS eu! MICROPEPLUS FULVUS ERICHSON eu x MICROPEPLUS MARIETTI DUVAL st! MICROPEPLUS LONGIPENNIS KRAATZ st! MICROPEPLUS PORCATUS (PAYKULL) eu x Staphylinidae PHLOEOCHARIS SUBTILISSIMA MANNERHEIM eu x 45! METOPSIA CLYPEATA (MÜLLER)! MEGARTHRUS DEPRESSUS (PAYKULL) eu x 53! MEGARTHRUS FRANZI SCHEERPELTZ st! MEGARTHRUS SINUATOCOLLIS (BOISDUVAL&LAC.) eu x 45, 53! MEGARTHRUS AFFINIS MILLER eu! MEGARTHRUS DENTICOLLIS (BECK) eu x MEGARTHRUS NITIDULUS KRAATZ eu MEGARTHRUS HEMIPTERUS (ILLIGER) eu x PROTEINUS OVALIS STEPHENS eu x 53! PROTEINUS CRENULATUS PANDELLE eu PROTEINUS LONGICORNIS DODERO st! PROTEINUS BRACHYPTERUS (FABRICIUS) eu x 45! PROTEINUS ATOMARIUS ERICHSON eu x PROTEINUS MACROPTERUS (GRAVENHORST) eu x EUSPHALERUM PALLENS (HEER) st x EUSPHALERUM ALPINUM (HEER) eu x EUSPHALERUM ANALE (ERICHSON) st x! EUSPHALERUM PALLIGERUM (KIESENWETTER) x EUSPHALERUM LONGIPENNE (ERICHSON) eu x EUSPHALERUM STRAMINEUM (KRAATZ) st x EUSPHALERUM MINUTUM (FABRICIUS) st x 45, 53! EUSPHALERUM ROBUSTUM HEER st x! EUSPHALERUM ABDOMINALE (GRAVENHORST) eu x 45! EUSPHALERUM LUTEUM (MARSHAM) eu x EUSPHALERUM MARSHAMI (FAUVEL) eu x 45! EUSPHALERUM SIGNATUM (MAERKEL) eu x 45! EUSPHALERUM LIMBATUM (ERICHSON) eu x 45! EUSPHALERUM PSEUDAUCUPARIAE (STRAND) eu x EUSPHALERUM RECTANGULUM (FAUVEL) eu x EUSPHALERUM RHODODENDRI (BAUDI) st EUSPHALERUM SORBI (GYLLENHAL) eu x 45 x EUSPHALERUM TORQUATUM (MARSHAM) eu x x EUSPHALERUM ATRUM (HEER) eu x 45! EUSPHALERUM FLORALE (PANZER) eu x x EUSPHALERUM KAHLENI ZANETTI! ACRULIA INFLATA (GYLLENHAL) st ACROLOCHA MINUTA (OLIVIER) eu x PHYLLODREPA NIGRA (GRAVENHORST) eu x 20